Alumni

	As a Putnam Fellow, Camilo Villouta focused on understanding the tradeoff between different survival strategies that trees employ with successful reproduction in the spring. Winter presents significant challenges to long lived perennial species, and in particular, to the reproductive buds that form before winter. Trees have developed different strategies for surviving winter and ensuring reproductive success.This work will add insight to how climate change may impact temperate woody species.
	Lindsay McCulloch was a NOAA Climate and Global Change postdoctoral fellow working in the Taylor lab. Lindsay’s work focuses on the role plant-microbial symbioses play in the biogeochemical cycling and function of tropical forests, and how these processes may affect global carbon and nutrient cycling.
	Lauren Church was a research assistant in the Taylor lab, and helped with sample processing, data analysis, as well as field work (her favorite part). Lauren completed her undergraduate thesis in the Taylor lab looking at the impacts of fertilizer treatments on soil bacterial communities, plant health and soil nutrients. Her research interests include global change biology, environmental justice, and human-environmental interactions.
	As a research assistant in the Friedman lab, Ellie Mendelson contributed to a variety of projects related to plant evolution and morphology. For her undergraduate thesis, Ellie studied the fruit developmental network in early diverging angiosperms Austrobaileya scandens and Illicium floridanum with the Ambrose Lab at the New York Botanical Garden.
	Hannah Adler, undergraduate student, Harvard College, Taylor Lab Spencer Carter, undergraduate student, Harvard College, Hopkins Lab Nick Daley, undergraduate student, Harvard College, Taylor Lab Isabel Eddy, undergraduate student, Harvard College, Taylor Lab Joseph Fadule, undergraduate student, Harvard College, Taylor Lab Jake Greer, undergraduate student, Harvard College, Hopkins Lab Andrew Ford, undergraduate student, Harvard College, Hopkins Lab Sabrina Freidus, undergraduate student, Harvard College, Hopkins Lab Kim Hartung, undergraduate student, Harvard College, Taylor Lab Peyton Jones, undergraduate student, Harvard College, Hopkins Lab Dorina Kodeanu, undergraduate student, Harvard College, Hopkins Lab Catherine Mailly, undergraduate student, Harvard College, Hopkins Lab

	Iñaki Hormaza is a Research Professor at IHSM la Mayora in Spain. As a Fulbright Scholar, he visited the Arboretum to study fruit development in the paw paw.
	Marwa Abdelal is a Co-Op Student from Northeastern University that worked in the Hopkins Lab.
	Wenying Liao was a Postdoctoral fellow in the Taylor Lab. Wenying combines statistical and theoretical modeling, greenhouse work, and policy analysis to study the global nitrogen cycle, focusing on the understanding of symbiotic nitrogen fixation in natural forests and the sustainable management of nitrogen use in agricultural landscapes.
	Arnold Arboretum Putnam Fellow and Visiting Scholar, Jake Grossman is a plant ecophysiologist with a strong interest in forest ecosystems and trees. Jake examines drought vulnerability and water use strategies in maples (Acer spp.) to analyze the potential effects of climate change on the Arboretum’s collections. Jake is an assistant professor at St. Oalf College.
	Cristina Ferrer Blanco is a PhD student at the IHSM La Mayora in Spain. She studys reproductive biology of an early divergent angiosperm, Asimina triloba (Pawpaw), mainly focused on its incompatibility system.
	Olivia Bronzo-Munich is an undergraduate student at Bowdoin College studying Biology. She is interested in plant-insect interactions and their role in plant and insect adaptation, speciation, and diversification. As a DaRin Butz intern, she worked in the Hopkins lab researching adaptation and speciation in the wildflower Phlox
	Cameron Chin (she/her) is an undergraduate at UMass Amherst studying environmental science and plant & soil science. Growing up in Boston, she’s enjoyed visiting the urban wilds and local parks to sketch the landscape and wildlife. She plans to go to graduate school to study plant/ microbe interactions under global change factors to work within the sphere of conservation. As an DaRin Butz intern in the Templer/ Hutyra Labs, she researched nutrient cycling in urban soil systems.
	Eden Fisher is an undergraduate at Harvard College interested in studying math, plants, rocks, public health, or some combination of the above. She grew up in Rhode Island and likes to spend as much of her time outside as possible. In the Friedman Lab as a DaRin Butz intern, Eden explored the connections between developmental, phylogenetic, and evolutionary biology.
	James Fortin grew up north of Boston and enjoys gardening, fishing, and exploring. He is a Biology major at a Williams College (at the opposite end of Massachusetts). After graduation, he plans to do fieldwork in evolutionary biology or ecology. In the Friedman Lab as a DaRin Butz Intern, James studied plant morphologies in the context of evolutionary diversification.
	William Huntley is an undergraduate at Bennington College in southern Vermont, studying biology with a focus in proteomics and plant biology. Originally from southwestern New Hampshire, he enjoys hiking, gardening, and being outside. He worked in the Moeglein lab researching leaf development as a DaRin Butz intern.
	Emily Lancaster is an undergraduate student at the University of Michigan studying plant biology. She is interested in plant adaptations and the affects of climate change on forest communities. As a DaRin Butz intern, Emily worked in the Villouta lab investigating winter physiology in plants.
	Joshua Murphy is an undergraduate student at The George Washington University studying Biology. He is interested in plant genetics and ethnobotany. As a DaRin Butz intern, he worked in the Hopkins lab researching adaptation and speciation in the Texas wildflower Phlox.
	Ben Tibbetts is an undergraduate student studying Biology at Case Western Reserve University. He is a violinist, plant lover, and particularly interested in the relationship between organisms and their environment. As a DaRin Butz intern he worked in the Taylor Lab investigating the effects of climate change on plants, as well as how plants can influence their surroundings in return.
	Antonio Serrato-Capuchina was a postdoctoral fellow in the Hopkins Lab interested in leveraging naturally hybridizing species complexes to uncover patterns in the development of reproductive isolation and better understand the evolution of species. Currently he is at Boston University.
	Jacob Suissa was a PhD Candidate in the Friedman Lab. He is an evolutionary biologist focusing on Ferns and Lycophytes. He takes a broad scale macroevolutionary approach as well as a small scale physiological approach to answer interesting questions about fern evolution. He is currently a postdoctoral fellow at Cornell University.
	Ben Goulet-Scott was a PhD Candidate in the Hopkins Lab. Ben is interested in adaptation and speciation in Phlox. Ben is now the higher education & lab coordinator at the Harvard Forest.
	A MEME Student in the Hopkins Lab, Bushra Shahid aimed to investigate whether there is a shift towards earlier and more autonomous selfing in sympatric populations of Phlox cuspidata compared to allopatric populations, constant with reinforcement favoring traits that reduce hybridization in sympatry.
	Daniel Buonaiuto was a PhD candidate in the Wolkovich and Holbrook Labs. He is interested in how plant communities are responding to global change, and how these responses are affecting the composition and function of North American ecosystems.
	Arturo Aguilar, undergraduate student, Harvard College, Taylor Lab. Kaitlyn DeGroot, undergraduate student, Harvard College, Friedman and Taylor Labs
	Kristel Schoonderwoerd was a PhD student in the Friedman Lab. Interested in many facets of evolutionary botany, she studied the diverse strategies temperate tree species use to overwinter embryonic leaves in resting buds and expand these leaves in the spring.
	Alexandria Pete is a PhD Student in the OEB department. She did a research rotation in the Friedman Lab. She is fascinated by the relationships that exist between evolutionary history and plant development. Specifically, she is interested in perisperm development, the theory of interparental conflict, and the double fertilization of angiosperms.
	Wendy Clement, a Visiting Scholar at the Arnold Arboretum and a Associate Professor at the College of New Jersey, studied floral traits using the living collections of Lonicera (honeysuckles) at the Arnold Arboretum. She is interested in using trait data, such as color, morphology, rewards, and floral scent to describe the evolutionary history of floral form in Lonicera and identify potential instances of shifts in pollination syndrome.
	Angie Diana was a research assistant in the Hopkins Lab.
	Al Kovaleski is a plant physiologist with a strong interest in understanding how woody plants adapt to freezing stresses during winter. As a Arnold Arboretum Putnam Fellow, Al focused on the Rosaceae family in the Arboretum’s living collections and examine cold hardiness, chilling response, and resumption of growth to recalibrate phenological models of budbreak.

	Elizabeth Spriggs was an Arnold Arboretum Putnam Fellow. As an evolutionary biologist with a passion for plant conservation, her research focused on the chestnuts and ashes (Castanea and Fraxinus) – iconic, but severely threatened, North American trees. Using phylogeography and genomics, she examined genetic diversity and population structure in relation to disease with an eye towards identifying individuals of conservation value.
	As a research assistant in the Hopkins Lab, Charlie Hale contributed to a variety of projects which combine field and greenhouse experiments with genomic analyses to study mechanisms of speciation in plants. Currently, he is a graduate student at Cornell University.
	Catherine Chamberlain was a PhD student in the Wolkovich and Holbrook Labs. She is interested in understanding how anthropogenic climate change affects plant communities and plant phenology.
	Samridhi Chaturvedi was a postdoctoral fellow in the Hopkins Lab. With an interest in the genomic basis of adaptation and speciation, she aimed to understand the genomic patterns of hybridization and introgression in Phlox species.
	Caio Guilherme Pereira is a postdoctoral fellow at MIT and a visiting fellow at the Arnold Arboretum. Caio aims to understand the molecular mechanisms of plant-environment interaction and discover how this interaction plays a role in the underlying growth pattern, composition and diversity of the plants.
	Steve Gougherty, PhD Student at Boston University, is developing a new framework to consider nutrient use efficiency (NUE) in plants beyond the traditional methods focused on vegetative tissues. As a Deland Award recipient, he measured the carbon and nitrogen costs of reproduction in samaras, a type of fruit. Samaras are green during seed development, suggesting active photosynthesis, but tend to lose their pigments as seeds mature.
	Aunnesha Bhowmick is an undergraduate student at University of California, Berkeley, studying Molecular Environmental Biology. She is passionate about climate change research especially through the lens of ecosystem ecology and biogeochemistry. As a DaRin Butz intern, she worked in the Taylor Lab where she analyzed remote sensing data to better understand the effects of climate change in the Arctic biome.
	Dylan Dubay is an undergraduate student at Goucher College studying Biology. He is interested in investigating adaptive genetic variance in hardwood trees and understanding forest ecology. As a Darin Butz Intern, he worked in the Grossman lab analyzing the potential effects of climate change on the Maple (Acer) genus.
	Jerry González-Cantoral is studying biology at University of San Carlos of Guatemala. His main research is focused on the influence of soil physicochemical properties on secondary metabolites secretion in flowers of the devil’s hand tree (Chiranthodendron pentadactylon). As a DaRin Butz intern, he worked in the Moeglein Lab exploring the complexities of leaf formation strategies and bud dormancy.
	Millie Harding studied Physical Geography at Durham University in the UK. Her research focused on land management techniques in the UK and the improvement of Payment for Ecosystem Services schemes in agriculture through interviews, choice and crop modelling. In the Taylor lab, she worked on spatial analysis of warming in the Arctic using remote sensing data.
	Erica Kirchhof is an undergraduate at Cornell University studying plant sciences with minors in climate change and inequality studies. She is interested in understanding the relationships between plants and the environment through the lens of physiology, ecosystem interactions, and global change. As a DaRin Butz intern, Erica worked in the Kovaleski Lab to explore the winter physiology of woody plants in the Arboretum’s living collections.
	Bea Mace is an undergraduate student at the College of Agriculture and Life Sciences at Cornell University studying Plant Science and Climate Change. She is interested in tree physiology, native landscape conservation, and the ecology of natural and cultivated woody landscapes. As a DaRin Butz intern, Bea worked in the Kovaleski Lab.
	Yanni Pappas is an undergraduate at Northeastern University studying Environmental Science and Political Science. He is interested in understanding the ways that human and natural systems interact, and how that knowledge can be used to help people live more sustainably with Earth’s environments. As a DaRin Butz intern, Yanni worked in the Templer Lab studying how air pollution affects urban ecosystems.
	As a Darin Butz Intern, Nidhi Vinod examined the impact of climate change on Maples (Acer genus) in the Grossman Lab.
	James Caven, undergraduate student, Harvard College, Hopkins Lab. Melissa Drake, undergraduate student, Harvard College, Hopkins Lab. Alex Paladino, undergraduate student, Harvard College, Hopkins Lab. Sophie Webster, undergraduate student, Harvard College, Hopkins Lab.

	Amelia Keyser-Gibson, Research Assistant, Friedman Lab and Weld Hill Labs, studied the reproductive development of conifers, as well as various other plant morphological questions.
	Rebecca Povilus, Postdoctoral Fellow, MIT and Visiting Fellow, Arnold Arboretum of Harvard University, is interested in the idea that molecular resources are important tools for connecting how changes at the gene and genome level affect developmental processes. She focuses on how evolutionary changes during the development of the egg-producing structure could give rise to the wide array of egg-sac morphologies in angiosperms.
	Kasia Zieminska, Arnold Arboretum Putnam Fellow, is interested in the role of anatomy on physiological function. Kasia focuses on the role of two wood tissues, fibers and parenchyma, in tree water storage.
	Paul Cervantes, undergraduate student, Harvard College, Friedman Lab. Skylah Reis, undergraduate student, Harvard College, Friedman Lab.
	As environmental engineer with interests in plant biology, Jie Yun is focused on solving practical environmental problems through the interdisciplinary research. She was a PhD student at MIT and Visiting Fellow of the Arnold Arboretum working in the Des Marais Lab.
	Barry Logan in collaboration with Dave Des Marais, examined divergent host spruce responses to parasitic dwarf mistletoe infection in the Arboretum. Barry is a Professor and Associate Dean of Bowdoin College and was a Visiting Scholar of the Arnold Arboretum of Harvard University.
	Natalia Pabón-Mora is interested in understanding the evolution of flower and fruit development and the morphological changes driving diversification of flowering plants. As a Jewett Prize recipient, she focuses on fruit diversity by comparing morphological and anatomical features and transcriptomic data of members of the Rubiaceae with distinct fruit types. Natalia is an Associate Professor at the University of Antioquia in Colombia and a Visiting Scholar of the Arnold Arboretum of Harvard University.

	Julia Marrs was a PhD Candidate at Boston University and Visiting Felllow of the Arnold Arboretum of Harvard University. Julia’s interests include terrestrial carbon cycling and remote sensing of vegetation. She investigated the link between the solar-induced chlorophyll fluorescence signal measured by tower and satellite platforms and plant physiological processes occurring at the leaf level.
	As a research assistant in the Friedman Lab, Laura Clerx studied early evolutionary thought, with a focus on biologists (especially botanists) who made contributions to evolutionary theory, both prior to and concurrent with Charles Darwin.
	Matt Farnitano was a research assistant in the Hopkins Lab. Starting Fall 2019, Matt is a graduate student at the University of Georgia.
	Liam Cleary is an undergraduate student at UMass Amherst studying plant and soil science. As a DaRin Butz Intern, Liam worked in the Friedman Lab studying Magnolia macrophylla’s relationship with honey bee pollinators.
	As a DaRin Butz Intern, Henry (Buster) Coe, an undergraduate at Oberlin College, worked in the Grossman Lab.
	Yiling Fang is an undergraduate at Oberlin College studying Biology. As a DaRin Butz Intern, she worked in the Holbrook Lab to understand cavitation resistance.
	Mayerlin Fischbach is an undergraduate student at the University of Central Florida, studying biology with a focus on plant science. As a DaRin Butz Intern, she worked to understand ecosystem resilience and plant-environment interactions in the Des Marais Lab.
	Fiona Harrigian is an undergraduate student at Wellesley College studying Biology. As a DaRin Butz Intern, she explored the connections between urbanization and plants and soils in the Templer and Hutyra Labs.
	Daniel Mindich is an undergraduate student at Harvard College studying Environmental Science and Public Policy. Danny was a DaRin Butz Intern in the in the Friedman Lab.
	Emma O’Donnell is an undergraduate student at UMass Amherst, majoring in biology. As a DaRin Butz Intern, she worked in the Hopkins Lab on the processes of speciation in Phlox.
	Dannielle Waugh is an undergraduate at the University of Central Florida, currently obtaining her bachelor’s degree in Biology with a minor in Environmental Sciences. As a DaRin Butz Intern, she worked in the Spriggs Lab studying the evolution and plant conservation of the chestnuts and ashes (Castanea and Fraxinus).
	Erin Wright is an undergraduate at Harvard College. As a DaRin Butz Intern, she worked in the Holbrook Lab.
	Shayla Salzman was a PhD Candidate in OEB and Fellow of the Arnold Arboretum. In the Hopkins and the Pierce Labs, Shayla focused on Cycads and their weevil pollinators and how scent variation leads to the species-specific mutualisms.
	In the Hopkins Lab, Franchesco Molina investigated how the genes involved in meiosis are locally adapted to different temperatures in Arabidopsis arenosa. Francheso was a PhD Student and Fellow of the Arnold Arboretum
	Dong Wang is a Sargent Award recipient and Assistant Professor at the University of Massachusetts, Amherst. He is interested in the strategies that long-lived woody legumes use to interact with rhizobia, the nitrogen-fixing bacteria the live in nodules on the roots. He examined the rhizobia present in root nodules at the Arboretum.
	As part of the Erasmus Mundus Master Program in Evolutionary Biology, Henry North conducted research in the Hopkins Lab for 6 months. Henry focused on the genetic mechanisms of pollination in Phlox.
	The research of Adam Roddy, postdoctoral fellow at Yale University, focuses on the physiological aspects of the evolution of flowering plants. As a Jewett Prize recipient, he examined 25 species of basal angiosperms in the Arboretum’s living collections, Adam measured physiological traits associated with water balance to examine the trade-offs between water transport, water storage, and the ability to maintain turgid and showy flowers.
	Harold Suarez Baron is a Deland Award recipient and PhD Candidate at the University of Antioquia in Colombia. He studied Aristolochia (dutchman’s pipe) in the Arboretum’s collections. In Aristolochia, the trichomes (hairs) found on flowers are important for attracting and retaining flies for pollination. Harold focuses on the genetic mechanisms underlying trichome development in this non-model plant.
	Andrea Brown, undergraduate student, Harvard College, Hopkins Lab.

	Marjorie Lundgren is a plant ecophysiologist. As a postdoctoral fellow at MIT and Arboretum Visiting Fellow, she focused on deconstructing the components of perenniality with an aim to inform crop improvement strategy.
	Juan M. Losada was a postdoctoral fellow in the Friedman, Holbrook and Leslie Labs. He is interested in the biochemical communication events between the male and female during pollination and fertilization. He focused on the evolution of the nourishing behavior of the perisperm and endosperm.
	Federico Roda is interested in understanding the functional mechanisms of environmental adaptation in plants. As postdoctoral fellow in the Hopkins Lab, Federico investigated the molecular basis of the evolution of reproductive isolation in the genus Phlox.
	Bridget Bickner studies biology at the University of Nebraska-Lincoln, where her research focuses on the evolutionary implications of sexual conflict and mating strategies in spiders. In the future, she plans to focus on microevolutionary processes. As a DaRin Butz Intern and under an American Society of Plant Biologists fellowship, Bridget worked in the Hopkins Lab to locate cis-regulatory promoter mutations related to Phlox drummondii flower color variation.
	DaRin Butz Intern Anna Blaustein is an undergraduate student at Bowdoin College where she studies Biology and Earth and Oceanographic Science. Anna worked in the Des Marais Lab investigating how CO2 levels affect resource allocation in cereal grain-like grasses.
	Emily Brown is an undergraduate student at Boston University studying Biology with a minor in Anthropology. As a DaRin Butz Intern, she worked on the project “Tree Anatomy and Function” in the Zieminska Lab.
	DaRin Butz Intern Matt Fertakos is an undergraduate studying biology at The College of New Jersey. His major interests include ecology, evolution, and conservation. Matt worked in the Spriggs Lab studying evolution and plant conservation of the severely threatened chestnut and ash trees.
	DaRin Butz Intern Anny Garcés Palacio is from Colombia, currently getting her undergraduate degree in Biology at Antioquia University. She is interested in Evo-Devo plants, evolutionary biology, organismic and embryology. Anny worked in the Friedman Lab exploring the process of double fertilization in Conifers.
	Zhe He is a recent graduate from Wheaton College with a chemistry major and biology minor. As a DaRin Butz Intern, Zhe worked in the Holbrook Lab to help characterize vascular xylem vulnerability to cavitation. In the Walhout lab at Wheaton, she studied the diffusion of rhodamine b in polyelectrolyte solutions.
	Marissa Lee is an undergraduate at Boston University and a DaRin Butz Intern. She worked on a project in the Templer and Hutyra Labs.
	Derek Schneider is a biology major at Amherst College. As a DaRin Butz Intern, he studied speciation in plants in the Hopkins Lab.
	Andrew Walter-McNeill is a undergraduate at Bowdoin College studying biology and computer science. As a DaRin Butz Intern, he investigated how certain crops respond to changes in their growth environment (especially those related to climate change) in the Des Marais Lab. He spent his spring semester studying the boundary between eucalypt forest and rainforest in northern Queensland.
	Isabelle Chuine is CNRS research director at the Centre d’Ecologie Fonctionnelle & Evolutive in Montpellier, France. As a Bullard Fellow, her project tackled the ongoing challenge of providing robust forecasts of climate change impacts on forest trees phenology. She tested key hypotheses on the regulation of cell seasonal activity using a series of experiments combining ecophysiology and biotracing methods.
	As part of the Aronia breeding program, Jonathan Mahoney, a Deland Award recipient and Master’s student at the University of Connecticut, investigated the mating systems and compatibility issues of intergeneric hybridization between Aronia and related taxa.
	Morgan Moeglein, a graduate student in the Edwards Lab at Yale University, studied the environmental and genetic factors governing leaf shape in the genus Viburnum utilizing the living collection of the Arnold Arboretum.
	Richard Li, postbac, Brandeis University, Wolkovich Lab. Sara Muchoney, undergraduate student, Boston University, Hopkins Lab. Nathan Oalican, undergraduate student, Harvard College, Friedman Lab.

	As a research associate in the Hopkins Lab, Sevan Suni investigated both the ecological factors and underlying genetic mechanisms that contribute to adaptation and speciation in Phlox populations. She is now an assistant professor at the University of San Francisco.
	Heather Briggs was a postdoctoral researcher in the Hopkins Lab. She focused on understanding pollination behavior of Battus philenor on Texas Phlox.
	The research of David Des Marais, Senior Fellow, focuses on how plants interact with the environment and the variation in these interactions between species. Understanding how plants adapt to the local environment can increase our ability to conserve plant populations. Dave is now an assistant professor at MIT.
	Rebecca Povilus was a PhD candidate in the Friedman Lab. She is interested in the idea that molecular resources are important tools for connecting how changes at the gene and genome level affect developmental processes. She focused on how evolutionary changes during the development of the egg-producing structure could give rise to the wide array of egg-sac morphologies in angiosperms.
	Ignacio Morales-Castilla was a research associate working in the the Wolkovich Lab. His research focused on the role of historical and evolutionary processes on current diversity patterns. He is interested in better understanding and predicting species and ecosystem vulnerability in a context of Global Change.
	Meng Li, a Ph.D student in the Chengdu Institute of Biology at the Chinese Academy of Sciences, is interested in the role of hybridization on species diversification. As part of his Hu Award, Li spent one year at the Arboretum to examine hybridization in Sorbus.
	Johan Arango earned his bachelor’s degree in Agroindustrial Engineering. He is currently pursuing a Master’s Degree in Sustainability and joined the Wolkovich Lab in the Fall of 2016. He was drawn to the lab because he is very interested in the effect that climate variation has on plants. He is very interested in societies’ reliance on plants and how a growing global population along with expanding markets will pose challenges and opportunities to achieve sustainability, especially in food systems.
	Wayne Daly graduated from Trinity College Dublin in 2016 with an MSc in biodiversity and conservation. He completed a study on geoxylic suffrutices, and a dissertation on salt marsh ecology and plant community compositions. As an intern in the Wolkovich Lab, he will learn more about the time effects and pressures on plants and plant phenology from various environmental factors and anthropogenic activities.
	Callin Switzer was a PhD Candidate in the Hopkins Lab. He is investigating bumblebee buzz pollination to understand how bees buzz and why there is variation in buzzing. Callin is now a a postdoctoral fellow in the eScience Institute at the University of Washington.
	Ashley Bang is an undergraduate at Brown University studying Geology-Biology. As a DaRin Butz Intern, she worked on the ‘Air Pollutions in Urban Ecosystems’ project in the Templer and Hutyra Labs. Originally from Hong Kong, Ashley is interested in studying the shift in ecosystem interactions and functions that have been caused by anthropogenic activities.
	Juliet Bramante is a Harvard undergraduate working in the Hopkins Lab. As a DaRin Butz Intern, she helped to identify cis regulatory changes responsible for color variation in Phlox drummondii.
	Vivien Chen is an undergraduate at Boston University studying Earth and Environmental Sciences with a minor in Biology. As a DaRin Butz Intern, she worked in the Holbrook Lab with Dr. Losada to determine whether a correlation exists with phloem size and ploidy in three species of white magnolias and in the Templer Lab and Hutyra Lab studying the effects climate change has on nonstructural carbohydrates in hardwood trees.
	Jessica Leslie is a recent graduate from UMass Amherst, where she was a Biology major with a certificate in Civic Engagement and Public Service. She is interested in plant-insect interactions, plant adaptations, and plant speciation. As a DaRin Butz Intern, she worked in the Hopkins Lab.
	Sruti Pandey is an undergraduate student at Boston University studying biology. As a DaRin Butz Intern, she worked with Dr. Losada in the Holbrook Lab to quantitatively study the vascular system of transgenic Nicotiana plants using confocal microscopy.
	Asa Peters, from Newton, Massachusetts is a sophomore at Connecticut College studying botany and music technology. He is interested in botany, ecology, and also ethnobotany. As a DaRin Butz Intern, Asa worked in the Wolkovich Lab.
	Emily Rosa is from Northern California currently getting her undergraduate degree at Sonoma State University in Environmental Studies and Planning with minors in Biology and Geography. As a DaRin Butz Intern, she worked in the Kasia Zieminska, Arnold Arboretum Putnam Fellow.
	Jack Smith is a Harvard undergraduate studying Environmental Science and Public Policy. As a DaRin Butz Intern, he worked in the Friedman Lab exploring plant morphology and development.
	Dan Flynn was a research associate in the Wolkovich Lab. He is interested in understanding the processes in which plant communities assemble and disassemble and how this will influence the ability of ecosystems to adapt to global climate change.
	Research associate and principal investigator Cam Webb studies the evolution and ecology of tropical plants, and is experimenting with new informatics and capacity-building approaches to plant inventory and taxonomy. Currently, Cam is collaborating with co-PI Sarah Mathews on a National Science Foundation funded project that focuses on the ecology, biogeography, and biodiversity informatics of trees in Indonesian forests.
	The research of Stacey Leicht Young, Arnold Arboretum Putnam Fellow, examines the ecological and reproductive strategies required for lianas (woody vines) to be successful in its environment. Utilizing the Arboretum’s Leventritt Shrub and Vine collection, Stacy compares and contrasts the functional traits of North American species with East Asian species growing in a common environment.
	The research interests of Cary Pirone, Arnold Arboretum Putnam Fellow, lay primarily in understanding how chemical signals mediate biological phenomena. She is exploring the complexities of pollination drops (ovular secretions) of several conifer species and ginkgo using biochemical and anatomical approaches.
	Jonathan Davies is an Associate Professor at McGill University in Canada. His work broadly addresses questions related to the distribution of biodiversity and the challenges posed to its conservation through recent changes to the environment.
	A PhD candidate from the University of Minnesota and a Deland Award Recipient, the research of Alexander Susko focuses on abiotic stress tolerance in Rhododendron. At the Arboretum, Alexander collected and sequenced deciduous azaleas originally collected across the geographic and environmental range to identify genetic targets of selection associated with differing environmental responses.
	A lab technician in the Wolkovich Lab, Tim Savas was involved in research focusing on community ecology in light of global climate change.
	Guan Chen, undergraduate student, Harvard College, Friedman Lab. Sasinat Chindapol, undergraduate student, Harvard College, Hopkins Lab. Sarah Gonzalez, undergraduate student, Harvard College, Hopkins Lab. Emily Guo, undergraduate student, Boston University, Hopkins Lab. Alice Linder, undergraduate student, Harvard College, Wolkovich Lab. Ashton Macfarlane, undergraduate student, Harvard College, Friedman Lab. Hanon McShea, undergraduate student, Harvard College, Friedman Lab. Christine Parkent, undergraduate student, Harvard College, Wolkovich Lab. Sam Schaffer-Morrison, undergraduate student, Wolkovich Lab. Elizabeth Stebbins, undergraduate student, Harvard College, Wolkovich Lab.

	Grace Yu, a research assistant in the Friedman Lab, was involved in projects studying angiosperm (flowering plant) reproductive and fertilization biology and embryology.
	Chase Mason, Arnold Arboretum Putnam Fellow, studies the evolution of leaf physical and chemical defenses in relation to the leaf economics spectrum (LES) across 16 woody genera. LES relates the initial investment of carbon during leaf development with the net carbon gain of that leaf. Plants employ various LES strategies which impacts their adaptability to environmental conditions and stresses.
	Rubén Sancho was a PhD student at the University of Zaragosa in Spain. He worked with Dave Des Marais to sequence the transcriptome of multiple Spanish species of Brachypodium, an important model grass species.
	Eric Goolsby is a post-doctoral researcher at Edwards Lab at Brown University. His research focuses on developing methods to examine the evolution of phenotypic responses to abiotic stress. The Arboretum collections allowed him to test his methods in a wide range of species.
	Jorge Lora is a Jewett Prize Recipient and a post-doctoral fellow with Professors Iñaki Hormaza and Maria Herrero at Experimental Research Stations – CSIC, Spain. Using members of the Arboretum’s Rosaceae collection, he compared ovule morphology from the earliest stages to maturity as well as the expression pattern of a gene thought to be important in ovule development.
	As a NSF post-doctoral fellow in Cynthia Weinig’s lab at the University of Wyoming, Rob Baker is investigating the relationship between anatomical features and water use efficiency in diverse cultivars of Brassica rapa.
	Jessica Savage is interested in understanding how seasonal changes in vascular activity influence flowering and carbon allocation. As an Arnold Arboretum Putnam Fellow, Jessica compared xylem and phloem anatomy, physiology and function in precocious flowering species (those that flower before the leaves emerge) with related species that flower later in the season.
	Jehane Samaha was a lab technician in the Wolkovich Lab. She examined functional traits and phenology of trees out in the field, in the lab, and in the greenhouse to understand how communities respond to global climate change.
	Elisabeth Forrestel was a postdoctoral fellow in the Wolkovich Lab. The focus of her research was to understand how an important economic crop, wine grapes, respond to climate change. She will study their responses both in the field and after manipulating their environment in growth chamber studies.
	Ben Ainsworth, undergraduate student, UMass, Boston, Hopkins Lab. Ryan Antolick, undergraduate student, Wheelock College, Wolkovich Lab. Jacob Barnett, Acton-Boxborough Regional High School teacher, Des Marais Lab. Lauren Brill, undergraduate student, Tufts University, Goolsby Lab. Aden Brown, undergraduate student, Tufts University, Hopkins Lab. Andrew Clark, undergraduate student, Harvard College, Hopkins Lab. Melissa DiTucci, undergraduate student, Mount Holyoke College, Hopkins Lab. Justin Dower, undergraduate student, Hopkins Lab. Harold Eyster, undergraduate student, Harvard College, Wolkovich Lab. Sally Gee, undergraduate student, Harvard College, Ettinger and Wolkovich Labs. Amna Jamshad, undergraduate student, University of Georgia, Mason Lab. Amna received a Zeller Summer Scholarship in Medicinal Botany from the Garden Club of America. Joe Kearney, undergraduate student, Harvard College, Hopkins Lab. Ari Korotkin undergraduate student, Harvard College, Wolkovich Lab. Michael LaScaleia, undergraduate student, Tufts University, Mason Lab. Evan Liu, undergraduate student, Harvard College, Hopkins Lab. Hanon McShea, undergraduate student, Harvard College, Hopkins Lab. Nicole Merrill, undergraduate student, Northeastern University, Wolkovich Lab. Sara Muchoney, undergraduate student, Boston University, Hopkins Lab. Cara O’Conner, undergraduate student, Harvard College, Wolkovich Lab. Tessa Pliakas, undergraduate student, Boston University, Mason Lab. Catherine Principe, postbac, Boston University, Wolkovich Lab. Luca Russo, undergraduate student, Boston University, Mason Lab. Corrinne Smith-Winterscheidt, undergraduate student, Tufts University, Mason Lab. Mark Tracy, undergraduate student, Harvard College, Mason Lab. Juliana Webber, undergraduate student, Boston University, Mason Lab. Adele Woodmansee, undergraduate student, Harvard College, Friedman Lab. Margaret Zahrah, undergraduate student, Tufts University, Hopkins Lab. Mona Zayed, undergraduate student, Boston University, Savage Lab.

Diana Bernal was a visiting fellow in the Hopkins Lab. She investigated the developmental mechanisms underlying variation of the reproductive mode in the Brachiaria.

Grey Monroe is a Ph.D. student at Colorado State University, Fort Collins that worked in the Des Marais Lab. Grey was awarded an Eden grant to come to the Arnold Arboretum for his research.

Emma Borjigin-Wang, Harvard undergraduate, Wolkovich Lab.
Chris Chen, Harvard undergraduate, Hopkins Lab.
Terilyn Chen, Harvard undergraduate, Wolkovich Lab.
Nakoa Farrant, Harvard undergraduate, Wolkovich Lab.
Silvia Golumbeanu, Harvard undergraduate, Ettinger and Savage Labs.
Sara Hamilton, undergraduate student, Bowdoin College, Des Marais Lab.
Risa Mcnellis, undergraduate student, Boston University, Ettinger Lab.
Jan Xue undergraduate student, University of British Columbia, Des Marais Lab. Jan was awarded an Eden grant to come to the Arnold Arboretum for her research.

Rubén Sancho was a PhD student at the University of Zaragosa in Spain. He worked with Dave Des Marais to sequence the transcriptome of multiple Spanish species of Brachypodium, an important model grass species.

Stuart Graham is a master’s student in the Erasmus Mundus Master Program in Evolutionary Biology. In the Hopkins Lab, he examined the behavior of pollinators that visit two sister species of Texas wildflower – Phlox drummondii and P. cuspidata.

A PhD candidate at the National University of Columbia, Marcela Serna is interested in understanding the role reproductive biology plays in the response of Magnolia to climate change to aid conservation efforts.

Barry Logan is a professor and associate dean at Bowdoin College. Collaborating with Bowdoin College undergraduate Sara Hamilton and Dave Des Marais, he examined the photosynthetic properties and drought tolerance of multiple accessions of Brachypodium sylvaticum, an important model grass species, which is also invasive in the Pacific Northwest.

A geneticist at the USDA Forest Service and a Sargent Award recipient, Andrew Groover performed RNA sequencing (RNA-seq) on cambium and wood forming tissues of diverse trees in the Arboretum. Potentially important regulatory genes expressed in these tissues will be identified and analyzed as a first step to understanding the evolution of woody growth.

Rosanne Healy, Sargent Award recipient and post-doctoral fellow at the University of Minnesota, focused on ectomycorrhizal fungi that are present on roots of trees in well-spaced habitats like the Arboretum. She sequenced, identified and compared the fungal partners on the roots of Oak trees present at the Arboretum with those at the Harvest Forest.

Conservation of threatened plant species is of great interest to Abby Hird. As a scientist for Botanic Gardens Conservation International–United States (BGCI-US), she collaborated with the Arnold Arboretum to implement the recommendations of the North American Collections Assessment. The goal is to preserve threatened taxa and increase genetic diversity by growing threatened plants in the living collection of the Arboretum and other botanical gardens.

Magaly Gutierrez is a Harvard undergraduate student that worked in the Wolkovich Lab.

Julia Paltseva was a summer research assistant in the Wolkovich Lab. She assisted in setting up lab and field experiments both at the Arboretum and at the Harvard Forest.

A recent graduate from the University of Connecticut, Kate Morozova was a Research Assistant in the Friedman Lab. One project that she was involved with was to examine the surface structure of Ginkgo biloba using scanning electron microscopy.

Ling Guo is a curator at the Beijing Botanic Garden and a Jewett Prize recipient. Utilizing the Arboretum’s Malus collection, she conducted research to improve the databases and knowledge of ornamental crabapples as a Registration Authority. Her database will focus on the flowers of Malus to improve the understanding of flowering time.

Utilizing the conifer collection, Guangyou Hao, Arnold Arboretum Putnam Fellow, studied the differences in the structural, physiological, and mechanical properties of water transport and xylem hydraulics between evergreen and deciduous conifers (which shed their leaves). Guangyou is now a Research Professor at the Institute of Applied Ecology at the Chinese Academy of Sciences.

Ursula King is a PhD student at the University of Connecticut, interested in the evolution of aquatic plants. Specifically, she is developing molecular markers to explore the population dynamics and reproductive biology of Najas flexilis (Hydrocharitaceae), a monoecious, freshwater annual, in which pollination is entirely underwater. As a visiting fellow in the Friedman Lab, Ursula learned techniques to observe ovule development in this species.

Harvard Forest E. C. Jeffrey Professor of Biology, Emeritus Barry Tomlinson has done extensive research on developmental morphology of pollen and cones in conifers. He collaborated with the Arboretum to investigate the composition and role of pollination drops in conifers.

Iñaki Hormaza from La Mayora Experimental Station in Malaga, Spain, is interested in flower and fruit development and pollen-pistil interactions. He spent time at the Arnold Arboretum studying the Rosaceae collection.

Sarah Mathews was a Sargent Fellow and principal investigator at the Arnold Arboretum. She is interested in plant phylogenetics and in the question of how changes in light-sensing systems have influenced the ability of plants to survive and diversify. The Mathews Lab uses phylogenetic, genetic, and comparative physiological approaches to explore the links between molecular and functional evolution in the phytochrome photoreceptor family.

A PhD Candidate at the University of Melbourne in Australia, Stephanie Conway is interested in the role of the shoot apical meristem in the evolution of shoot architecture. As a visiting fellow in the Friedman Lab, she focused on shoot apical meristems utilizing the extensive collection of gymnosperms at the Arboretum.

Julien Bachelier was a post-doctoral fellow in the Friedman Lab. His work focuses on the evolution of seed plants, particularly in the origin of flowering plants and the evolutionary development of flower and carpel. Julien’s thesis investigations compared the structure and development of flowers in closely related families such the Anacardiaceae and the Burseraceae.

A visiting scholar in the Friedman Lab, Biao Jin, Associate Professor at Yangzhou University, is interested in exploring the evolution of reproductive biology from gymosperms to angiosperms.

Jorge Lora is a Jewett Prize Recipient and a post-doctoral fellow with Professors Iñaki Hormaza and Maria Herrero at Experimental Research Stations – CSIC, Spain. Using members of the Arboretum’s Rosaceae collection, he compared ovule morphology from the earliest stages to maturity as well as the expression pattern of a gene thought to be important in ovule development.

The research of Francesca Secchi, a postdoctoral researcher in the Zwieniecki Lab, aims at understanding the principles of the biological function of vascular systems in plants. Her current research is focused on embolism formation and refilling using molecular, physiological, and biophysical approaches.

Arnold Arboretum Sargent Fellow Maciej Zwieniecki addresses the structure, surface chemistry, and mechanical properties of the network of fluid conduits in plants. The Zwieniecki Lab studies the specialized cells that distribute water, solutes, and energy over long distances, as well as how the terminal exchange surfaces (roots and leaves) interface with the environment.

Stuart Davies is the director of the Center for Tropical Forest Science at the Smithsonian Tropical Research Institute. He is interested in the ecology and evolution of tropical rainforests. Using a global network of large-scale long-term forest monitoring plots, he is investigating how environmental variation and change affect the diversity and dynamics of forests.

The research of Noeline Morrissey, a master’s student in the Friedman Lab, focused on trichome morphology and development. Comparing the multiple forms of trichomes in hops (Humulus lupulus) and related genera should shed light on the evolution of these highly specialized structures.

Erica Fadon is a PhD student in the Herrero Lab at the Pomology Department of the Aula Dei Experimental Station–CSIC, Spain. Using the Arboretum’s living collection, Erica closely examined the development of sweet cherry flowers (Prunus avium) in order to determine which stage the flowers undergo winter dormancy.

Hugh McAllister, honorary lecturer at the University of Liverpool, is interested in examining the evolutionary relationships of species within genera. By comparing chromosome numbers, he seeks to clarify the relationships between diploid and polyploid species. Visits to the Arnold Arboretum and the Harvard University Herbaria have led to the publication of monographs on genera Betula and Sorbus.

Elizabeth Ryan is an undergraduate student at Brown University working with Dov Sax. As an Arboretum Visiting Fellow, she worked with Michael Dosmann to research plant distributions in the face of climate change.

Kang Min Ngo is a research assistant for the CTFS. Based in Singapore, she is interested in understanding the role of forests in a changing climate, particularly through carbon sequestration.

Jessica Savage is a post-doctoral fellow in the Holbrook Lab at Harvard University and collaborated with Maciej Zwieniecki to study phloem loading in trees and vines. Her background is in plant hydraulics and ecophysiology.

From the University of Melbourne, Australia, the research of Denise Johnstone focuses on the physiological mechanisms involved in drought response in urban trees. As a visiting scholar in the Zwieniecki Lab, she examined xylem physiology during drought stress.

As a graduate in the Diggle Lab at the University of Colorado, Rob Baker focused on the evolution of shoot architecture by examining the molecular developmental pathways within and among populations, where genetic divergence, adaptation, and speciation occur. Specifically, he examined natural variation in shoot architecture patterning in Mimulus gattatus. Currently, he is a post-doc in Cynthia Weinig’s lab at the University of Wyoming.

David Kenfack is the coordinator for the Center for Tropical Forest Science-Smithsonian Institution Global Earth Observatory (CTFS–SIGEO) Africa Program. His primary interests are the use of plant systematics to explore species limits and exploring the ecological processes that explain tropical forest dynamics in Africa.

Elena Kramer is a professor in the Department of Organismic and Evolutionary Biology at Harvard University. The Kramer Lab is interested in the evolution of plant developmental genetics with a particular focus on floral evolution. Elena spent her sabbatical at the Weld Hill Research Building of the Arnold Arboretum, primarily writing but also taking advantage of the Arboretum’s microscopes and beautiful surroundings.

Brian Morgan was an Arnold Arboretum Putnam Fellow, and his research focused on the use of geographic information systems (GIS) technology as a management and decision-making tool for public gardens. Brian created a GIS platform for the Arnold Arboretum based on the ArcGIS Public Garden Data Model, and created a web-based application for performing collections research. He is the director of the Alliance for Public Gardens GIS.

Erin Kurten was a post-doctoral fellow working with Stuart Davies and CTFS-AA. Erin’s work focuses on tropical plant community ecology and the interactions which underlie community assembly processes and biogeographic patterns. With CTFS, Erin investigated how tree species adapt their phenology in response to drought stress in the seasonally-dry tropical forests of Southeast Asia.

Juan M. Losada is a graduate student in the Pomology Department of the Aula Dei Experimental Station–CSIC, Spain. He is interested in the events between pollination and fertilization, primarily the role of arabinogalactan proteins in pollen-pistil interactions in Malus. During his stay at the Arboretum, he worked with Ned Friedman to expand his dissertation studies to include Magnolia and Stewartia.

Iñaki Hormaza from La Mayora Experimental Station in Malaga, Spain, is interested in flower and fruit development and pollen-pistil interactions. He spent his summer at the Arnold Arboretum studying the Rosaceae collection.

Scherbatskoy was a Deland Award recipient working in the Friedman Lab with Julien Bachelier. In an effort to shed light on the evolution of female gametophyte development, the research focused on the comparative morphology of female gametophytes in diverse conifers.

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