All Scientists
About Alumni
2024

Alaina Bisson
2024 Sinnott Award
Research Assistant, OEB, Taylor Lab, Harvard University

Andrea Berardi
Research Associate, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University

Andrew Cameron
Research Assistant, OEB, Hopkins Lab

Anna Feller
Swiss National Science Foundation Postdoctoral Fellow, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University
Anya Ghai
undergraduate student, Northeastern University, Hopkins Lab

Cecilia (Ceci) Prada Cordero
Postdoctoral Fellow, OEB, Taylor Lab
Fellow of the Arnold Arboretum of Harvard University
Cheng-Chiang Wu
Assistant Professor, Framingham State University
Visiting Fellow, Arnold Arboretum of Harvard University
Irene Ren
undergraduate student, Northeastern University, Hopkins Lab
Isabella Cao
Undergraduate student, Harvard College, Taylor Lab
Luke Zehr
postgraduate student, Taylor Lab
Michael Fairley
Undergraduate student, Harvard College, Taylor Lab
Michael Fairly
undergraduate student, Harvard College, Taylor lab
2023

Austin Garner
Graduate Student, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University

Camilo Villouta
2021-2023 Putnam Fellow
Ellie Mendelson
Research Assistant, Friedman and Weld Hill Labs

Iñaki Hormaza
Research Professor, La Mayora Experimental Station, Spain
Visiting Scholar, Arnold Arboretum of Harvard University

Lauren Church
Research Assistant, OEB, Taylor Lab

Lindsay McCulloch
NOAA Climate and Global Change Postdoctoral Fellow
Fellow of the Arnold Arboretum of Harvard University

Megan Wilcots
2023 Global Change Fellow
Rohan Prabhu
undergraduate student, Hopkins lab
Undergraduate Student Interns
2022

Alexandria Pete
Graduate Student, OEB, Friedman and Kramer Labs
Angie Diana
Research Assistant, OEB, Hopkins Lab

Antonio Serrato-Capuchina
Postdoctoral Fellow, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University

Ben Goulet-Scott
Graduate Student, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University

Bushra Shahid
MEME Student, OEB, Hopkins Lab

Cristina Ferrer Blanco
PhD Student, IHSM La Mayora in Spain
Visiting Fellow, Arnold Arboretum of Harvard University

Daniel Buonaiuto
Graduate Student, OEB, Wolkovich and Holbrook Labs
Fellow of the Arnold Arboretum of Harvard University
Elizabeth Spriggs
2017-2019 Putnam Fellow
- Spriggs EL*, Fertakos ME. 2021. Evolution of Castanea in North America: restriction-site-associated DNA sequencing and ecological modeling reveal a history of radiation, range shifts, and disease. American Journal of Botany 108: 1692-704. Abstract

Jacob Suissa
Graduate Student, OEB, Friedman Lab
Fellow of the Arnold Arboretum of Harvard University

Jake Grossman
2018-2020 Putnam Fellow
- Grossman JJ*. 2023. Phenological physiology: seasonal patterns of plant stress tolerance in a changing climate. New Phytologist 237: 1
- Grossman JJ*. 2021. Evidence of constrained divergence and conservatism in climatic niches of the temperate Maples (Acer L.). Forests 12(5): 535. Abstract
- Kovaleski AP*, Grossman JJ*. 2021. Standardization of electrolyte leakage data and a novel liquid nitrogen control improve measurements of cold hardiness in woody tissue. Plant Methods 17: 53. (APK and JJG contributed equally) Abstract
Marwa Abdelal
Postgraduate Student, Northeastern University

Morgan Moeglein
2021-2022 Putnam Fellow
Assistant Professor, Norwich University
Visiting Scholar, Arnold Arboretum of Harvard University
Wendy Clement
Associate Professor, College of New Jersey
Visiting Scholar, Arnold Arboretum of Harvard University
Wenying Liao
Postdoctoral Fellow, OEB, Taylor Lab
2021

Al Kovaleski
2019-2021 Putnam Fellow
- Kovaleski AP*. 2022. Woody species do not differ in dormancy progression: Differences in time to budbreak due to forcing and cold hardiness. Proceedings of the National Academy of Sciences 119: e2112250119. Abstract
- Kovaleski AP*, Grossman JJ*. 2021. Standardization of electrolyte leakage data and a novel liquid nitrogen control improve measurements of cold hardiness in woody tissue. Plant Methods 17: 53. (APK and JJG contributed equally) Abstract

Caio Guilherme Pereira
Postdoctoral Fellow, MIT
Visiting Fellow, Arnold Arboretum of Harvard University

Catherine Chamberlain
Graduate Student, OEB, Wolkovich and Holbrook Labs
Fellow of the Arnold Arboretum of Harvard University

Charlie Hale

Kristel Schoonderwoerd
Graduate Student, OEB, Friedman Lab
Fellow of the Arnold Arboretum of Harvard University

Samridhi Chaturvedi
Postdoctoral Fellow, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University

Steve Gougherty
2019 Deland Award
PhD Student, Boston University
2020

Amelia Keyser-Gibson
Research Assistant, Friedman and Weld Hill Labs

Barry Logan
Professor and Associate Dean of Bowdoin College
Visiting Scholar, Arnold Arboretum of Harvard University.

Jake Grossman
2018-2020 Putnam Fellow
- Grossman JJ*. 2023. Phenological physiology: seasonal patterns of plant stress tolerance in a changing climate. New Phytologist 237: 1
- Grossman JJ*. 2021. Evidence of constrained divergence and conservatism in climatic niches of the temperate Maples (Acer L.). Forests 12(5): 535. Abstract
- Kovaleski AP*, Grossman JJ*. 2021. Standardization of electrolyte leakage data and a novel liquid nitrogen control improve measurements of cold hardiness in woody tissue. Plant Methods 17: 53. (APK and JJG contributed equally) Abstract

Jie Yun
Graduate Student, MIT
Visiting Fellow, Arnold Arboretum of Harvard University

Kasia Zieminska
2016-2018 Putnam Fellow
- Ziemińska K*. 2023. The role of imperforate tracheary elements and narrow vessels in wood capacitance of angiosperm trees. IAWA Journal 1(aop):1–14. Abstract
- Ziemińska K*, Rosa E, Gleason SM, Holbrook NM. Wood day capacitance is related to water content, wood density, and anatomy across 30 temperate tree species. Plant Cell Environ. 43: 3048– 3067.

Natalia Pabón-Mora
2018 Jewett Prize
Associate Professor, University of Antioquia in Colombia
Visiting Scholar, Arnold Arboretum of Harvard University
- Suárez-Baron H, Alzate JF, Ambrose BA, Pelaz S, González F, Pabón-Mora N. 2023. Comparative morphoanatomy and transcriptomic analyses reveal key factors controlling floral trichome development in Aristolochia (Aristolochiaceae). Journal of Experimental Botany74(21):6588–6607. Abstract

Rebecca Povilus
Postdoctoral Fellow, MIT
Visiting Fellow, Arnold Arboretum of Harvard University
Wayne Daly
Postgraduate student, Wolkovich Lab
2019
Adam Roddy
2016 Jewett Prize
Post-doctoral fellow, Yale University
Visiting Fellow, Arnold Arboretum of Harvard University
Elizabeth Spriggs
2017-2019 Putnam Fellow
- Spriggs EL*, Fertakos ME. 2021. Evolution of Castanea in North America: restriction-site-associated DNA sequencing and ecological modeling reveal a history of radiation, range shifts, and disease. American Journal of Botany 108: 1692-704. Abstract

Franchesco Molina
Graduate Student, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University

Harold Suarez Baron
2018 Deland Award
PhD Candidate, University of Antioquia in Colombia
Visiting Fellow, Arnold Arboretum of Harvard University
- Suárez-Baron H, Alzate JF, Ambrose BA, Pelaz S, González F, Pabón-Mora N. 2023. Comparative morphoanatomy and transcriptomic analyses reveal key factors controlling floral trichome development in Aristolochia (Aristolochiaceae). Journal of Experimental Botany74(21):6588–6607. Abstract
Henry North
Graduate Student, OEB, Hopkins Lab

Julia Marrs
Graduate Student, Boston University
Visiting Fellow, Arnold Arboretum of Harvard University
Laura Clerx
Research Assistant, Friedman Lab
Matt Farnitano
Research Assistant, OEB, Hopkins Lab
Shayla Salzman
Graduate Student, OEB, Hopkins and Pierce Labs
Fellow of the Arnold Arboretum of Harvard University
2018
Diana Bernal
Postdoctoral Fellow, Northeastern University
Visiting Fellow, OEB, Hopkins Lab
Federico Roda
Postdoctoral Fellow, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University
Heather Briggs
Postdoctoral Fellow, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University
Isabelle Chuine
CNRS Research Director, Centre d’Ecologie Fonctionnelle & Evolutive
Bullard Fellow, Harvard Forest

Jessica Savage
2014-2016 Putnam Fellow
2017 Sinnott Award
2023 Jewett Prize
Associate Professor, University of Minnesota, Duluth
Visiting Scholar, Arnold Arboretum of Harvard University
- Savage JA, Kiecker T, McMann N, Park D, Rothendler M, Mosher K. 2022. Leaf out time correlates with wood anatomy across large geographic scales and within local communities. New Phytologist 235: 953-64. Abstract

Juan M. Losada
Postdoctoral Fellow, Friedman, Holbrook, and Leslie Labs

Kasia Zieminska
2016-2018 Putnam Fellow
- Ziemińska K*. 2023. The role of imperforate tracheary elements and narrow vessels in wood capacitance of angiosperm trees. IAWA Journal 1(aop):1–14. Abstract
- Ziemińska K*, Rosa E, Gleason SM, Holbrook NM. Wood day capacitance is related to water content, wood density, and anatomy across 30 temperate tree species. Plant Cell Environ. 43: 3048– 3067.

Marjorie Lundgren
Postdoctoral Fellow, MIT
Visiting Fellow, Arnold Arboretum of Harvard University

Morgan Moeglein
Graduate Student, Yale University
Visiting Fellow, Arnold Arboretum of Harvard University
Sevan Suni
Postdoctoral Fellow, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University
2017
Callin Switzer
Graduate Student, OEB, Hopkins Lab
Fellow of the Arnold Arboretum of Harvard University
Cam Webb
Research Associate, Arnold Arboretum

David Des Marais
Research Associate, OEB
Senior Fellow of the Arnold Arboretum of Harvard University
Elizabeth Wolkovich
Associate Professor, University of British Columbia, Canada
Visiting Scholar, OEB, Harvard University
Ignacio Morales-Castilla
Research Associate, OEB, Wolkovich Lab
Fellow of the Arnold Arboretum of Harvard University

Jack Smith
Harvard University – 2019
2017 DaRin Butz Intern in the Friedman Lab
Johan Arango
Master's Student, Harvard Extension School, Wolkovich Lab
Jonathan Davies
Associate Professor, McGill University
Visiting Scholar, Arnold Arboretum of Harvard University

Jonathan Mahoney
2016 Deland Award
Master’s student, University of Connecticut
Visiting Fellow, Arnold Arboretum of Harvard University
- Mahoney JD, Brand MH. 2021. Pre- and postzygotic barriers associated with intergeneric hybridization between Aronia melanocarpa (Michx.) Elliott x Pyrus communis L. and ×Sorbaronia dippelii (Zabel) CK Schneid. x Pyrus communis. HortScience 56(2): 177-184. Abstract

Meng Li
2016 Hu Student/Postdoctoral Exchange Award
Ph.D student, Chengdu Institute of Biology, Chinese Academy of Sciences
Visiting Fellow, Arnold Arboretum of Harvard University
- Li M, Ohi-Toma T, Gao Y-D, Xu B, Zhu Z-M, Ju W-B, Gao X-F. 2017. Molecular phylogenetics and historical biogeography of Sorbus sensu stricto (Rosaceae). Molecular Phylogenetics and Evolution 111: 76-86.

Rebecca Povilus
Graduate Student, OEB, Friedman Lab
Fellow of the Arnold Arboretum of Harvard University
Rob Baker
NSF Postdoctoral Fellow, University of Wyoming
Visiting Fellow, Arnold Arboretum of Harvard University
2016

Chase Mason
2015-2016 Putnam Fellow
- De La Pascua DR, Smith‐Winterscheidt C, Dowell JA, Goolsby EW, Mason CM*. 2020. Evolutionary trade‐offs in the chemical defense of floral and fruit tissues across genus Cornus. Appl Plant Sci. 107(9):1260–73.
- Mason CM*, LaScaleia MC, De La Pascua DR, Monroe JG, Goolsby EW. 2020. Learning from dynamic traits: Seasonal shifts yield insights into ecophysiological trade-offs across scales from macroevolutionary to intraindividual. Int J Plant Sci. 181: 88-102.
Dan Flynn
Research Associate, OEB, Wolkovich Lab
Elisabeth Forrestel
Postdoctoral Fellow, OEB, Wolkovich Lab
Eric Goolsby
Postdoctoral Fellow, Yale University
Visiting Fellow, Arnold Arboretum of Harvard University
Grace Yu
Research Assistant, Friedman Lab
Jehane Samaha
Research Assistant, OEB, Wolkovich Lab
Jessica Savage
2014-2016 Putnam Fellow
- Savage JA*. 2019. A temporal shift in resource allocation facilitates flowering before leaf out and spring vessel maturation in precocious species. American Journal of Botany 106(1):113–22.
- Savage JA*, Beecher SD, Clerx L, Gersony JT, Knoblauch J, Losada JM, Jensen KH, Knoblauch M, Holbrook NM. 2017. Maintenance of carbohydrate transport in tall trees. Nature Plants 3: 965-972

Jorge Lora
2012 Jewett Prize
Post-doctoral Fellow, Experimental Research Stations (CSIC), Spain
Visiting Fellow, Arnold Arboretum of Harvard University
- Losada JM, Hormaza JI, Lora J. 2017. Pollen–pistil interaction in pawpaw (Asimina triloba), the northernmost species of the mainly tropical family Annonaceae. American Journal of Botany 104:1891–1903
Rubén Sancho
Graduate Student, University of Zaragosa
Tim Savas
Research Assistant, OEB, Wolkovich Lab
2015
Grey Monroe
Graduate Student, Colorado State University, Fort Collins
Marcela Serna
Graduate Student, National University of Colombia
Visiting Fellow, Arnold Arboretum of Harvard University
Stuart Graham
Erasmus Mundus Graduate Student, OEB, Hopkins Lab
2014

Ailene Ettinger
2014 Putnam Fellow
- Wolkovich EM, Chamberlain CJ, Buonaiuto DM, Ettinger AK, Morales-Castilla I. 2022. Integrating experiments to predict interactive cue effects on spring phenology with warming. New Phytologist 235: 1719-1728. Abstract
- Ettinger AK*, Buonaiuto DM, Chamberlain CJ, Morales-Castilla I, Wolkovich EM. 2021. Spatial and temporal shifts in photoperiod with climate change. New Phytologist 230: 462-474. Abstract
- Ettinger AK*, Chamberlain CJ, Morales-Castilla I, Buonaiuto DM, Flynn DFB, Savas T, Samaha JA, Wolkovich EM. 2020. Winter temperatures predominate in spring phenological responses to warming. Nature Climate Change.
- Ettinger AK*, Gee S, Wolkovich EM. Phenological sequences: how early-season events define those that follow. American Journal of Botany. 105: 1-1010(12):1137–42.

Andrew Groover
2014 Sargent Award
Geneticist, USDA Forest Service
Visiting Scholar, Arnold Arboretum of Harvard University

Guangyou Hao
2012-2014 Putnam Fellow
- Hao G-Y*, Lucero ME, Sanderson SC, Zacharias EH, Holbrook NM. 2013. Polyploidy enhances the occupation of heterogeneous environments through hydraulic related trade-offs in Atriplex canescens (Chenopodiaceae). New Phytologist 197: 970–8.
- Hao G-Y*, Wang A-Y, Sack L, Goldstein G, Cao K-F. 2013. Is hemiepiphytism an adaptation to high irradiance? Testing seedling responses to light levels and drought in hemiepiphytic and non-hemiepiphytic Ficus. Physiologia Plantarum 148: 74-86.
- Hao G-Y*, Wheeler JK, Holbrook NM, Goldstein G. 2013. Investigating xylem embolism formation, refilling and water storage in tree trunks using frequency domain reflectometry. Journal of Experimental Botany 64: 2321-2332.
Kate Morozova
Research Assistant, OEB, Friedman Lab

Ling Guo
2014 Jewett Prize
Curator, Beijing Botanic Garden
Visiting Scholar, Arnold Arboretum of Harvard University

Rosanne Healy
2013 Sargent Award
Post-doctoral Fellow, University of Minnesota
Visiting Fellow, Arnold Arboretum of Harvard University
- Healy RA, Zurier H, Bonito G, Smith ME, Pfister DH. 2016. Mycorrhizal detection of native and non-native truffles in a historic arboretum and the discovery of a new North American species, Tuber arnoldianum sp. nov. Mycorrhiza 26: 781-792.
- Leicht-Young SA, Healy R, Del Tredici P. 2016. Observation of mycorrhizal colonization in roots in natural populations of Celastrus orbiculatus Thunb. The Journal of the Torrey Botanical Society 143: 322-324.
- Shao D, Tang S, Healy RA, Imerman PM, Schrunk DE, Rumbeiha WK. 2016. A novel orellanine containing mushroom Cortinarius armillatus. Toxicon 114: 65-74.
Ursula King
Graduate Student, University of Connecticut
Visiting Fellow, Arnold Arboretum of Harvard University
2013
Biao Jin
Associate Professor, Yangzhou University
Visiting Scholar, Arnold Arboretum of Harvard University

Cary Pirone
2011-2013 Putnam Fellow
- Prior N, Little SA, Boyes I, Griffith P, Husby C, Pirone-Davies C*, Stevenson DW, Tomlinson PB, von Aderkas P. 2019. Complex reproductive secretions occur in all extant gymnosperm lineages: a proteomic survey of gymnosperm pollination drops. Plant Reprod. 32: 153-166.
- Nepi M, Little S, Guarnieri M, Nocentini D, Prior N, Gill J, Barry Tomlinson P, Ickert-Bond SM, Pirone C*, Pacini E, von Aderkas P. 2017. Phylogenetic and functional signals in gymnosperm ovular secretions. Annals of Botany 120: 923-936.
- Pirone-Davies C*, Prior N, Von Aderkas P, Smith D, Hardie D, Friedman WE, Mathews S. 2016. Insights from the pollination drop proteome and the ovule transcriptome of Cephalotaxus at the time of pollination drop production. Annuals of Botany 117: 973-984.
- Prior N, Little SA, Pirone C*, Gill JE, Smith D, Han J, Hardie D, O’Leary SJB, Wagner RE, Cross T, Coulter A, Borchers C, Olafson RW, and von Aderkas P. 2013. Application of proteomics to the study of pollination drops. Applications in Plant Science 1: 1300008.

Iñaki Hormaza
Research Professor, La Mayora Experimental Station, Spain
Visiting Scholar, Arnold Arboretum of Harvard University

Jorge Lora
2012 Jewett Prize
Post-doctoral Fellow, Experimental Research Stations (CSIC), Spain
Visiting Fellow, Arnold Arboretum of Harvard University
- Losada JM, Hormaza JI, Lora J. 2017. Pollen–pistil interaction in pawpaw (Asimina triloba), the northernmost species of the mainly tropical family Annonaceae. American Journal of Botany 104:1891–1903
Julien Bachelier
Postdoctoral Fellow, OEB, Friedman Lab
Sarah Mathews
Sargent Fellow, Arnold Arboretum of Harvard University

Stacey Leicht Young
2013 Putnam Fellow
- Leicht-Young SA*, Healy R, Del Tredici P. 2016. Observation of mycorrhizal colonization in roots in natural populations of Celastrus orbiculatus Thunb. The Journal of the Torrey Botanical Society 143: 322-324.
Stephanie Conway
Graduate Student, University of Melbourne, Australia
Visiting Fellow, Arnold Arboretum of Harvard University
Stuart Davies
Director, Center for Tropical Forest Science, Smithsonian Tropical Research Institute
2012
Denise Johnstone
Faculty, University of Melbourne, Australia
Elizabeth Ryan
Undergraduate Student, Brown University
Visiting Fellow, Arnold Arboretum of Harvard University
Erica Fadon
Graduate Student, Aula Dei Experimental Station–CSIC, Spain
Visiting Fellow, Arnold Arboretum of Harvard University
Francesca Secchi
Postdoctoral Fellow, AA, Zwieniecki Lab

Hugh McAllister
2012 Sargent Award
Honorary Lecturer, University of Liverpool
Visiting Scholar, Arnold Arboretum of Harvard University

Juan M. Losada
2011 Deland Award
PhD Candidate, Aula Dei Experimental Station–CSIC, Spain
Visiting Fellow, Arnold Arboretum of Harvard University
Kang Min Ngo
Maciej Zwieniecki
Sargent Fellow, Arnold Arboretum of Harvard University
Noeline Morrissey
Graduate Student, Harvard Extension School, Friedman Lab
2011
Brian Morgan
2010-2011 Putnam Fellow
David Kenfack
Coordinator, Center for Tropical Forest Science-Smithsonian Institution Global Earth Observatory (CTFS–SIGEO) Africa Program
E. Scherbatskoy
2011 Deland Award
Undergraduate Student, University of Colorado, Boulder
Erin Kurten
Postdoctoral Fellow, AA, Davies Lab

Iñaki Hormaza
Research Professor, La Mayora Experimental Station, Spain
Visiting Scholar, Arnold Arboretum of Harvard University
Rob Baker
Graduate Student, University of Colorado, Boulder
Visiting Fellow, Arnold Arboretum of Harvard University
2010
Abby Hird
2008-2010 Putnam Fellow
2009

Jennifer Winther
2007-2009 Putnam Fellow
2007

Richard Primack
2006-2007 Putnam Fellow
2006

Eric Hsu
2005-2006 Putnam Fellow
2005

Nina Theis
2004-2005 Putnam Fellow

Tim Brodribb
2005 Putnam Fellow
2004

Sonali Saha
2002-2004 Putnam Fellow
2003

Lawren Sack
2002-2003 Putnam Fellow

Sonia Uyterhoeven
2002-2003 Putnam Fellow
2002

Michael Dosmann
2000-2002 Putnam Fellow
Keeper of Living Collections
Arnold Arboretum of Harvard University

Youngdong Kim
2001-2002 Putnam Fellow
2001

Donglin Zhang
2001 Putnam Fellow
1999

Jianhua Li
1998-1999 Putnam Fellow
1998
Katherine Gould Mathews
1997-1998 Putnam Fellow
Keiko Satoh
1997-1998 Putnam Fellow
1996

Jinshuang Ma
1995-1996 Putnam Fellow

Kim Tripp
1994-1996 Putnam Fellow
1994

David Ackerly
1994 Putnam Fellow
1992

Jun Wen
1991-1992 Putnam Fellow
1991

Michael Dirr
1991 Putnam Fellow
1990

Elizabeth Kellogg
1989-1990 Putnam Fellow

Richard Schulhof
1989-1990 Putnam Fellow

Tokushiro Takaso
1988-1990 Putnam Fellow
1989

Jessica Gurevitch
1989 Putnam Fellow